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Horde - Horde Groupware is a free, enterprise ready, browser based collaboration suite.
Roundcube - Web-based IMAP email client
Mailo - Mailo is an email client where you can send and receive emails to and from anyone with an email address.
Mailpile - Mailpile is a modern, fast web-mail client with user-friendly encryption and privacy features.
Rainloop - RainLoop is a web based email client.
HEY - Email at its best, new from Basecamp.